Form and Substance of Expression

[Hjelmslev] used the term expression for functional structures, which would also have to be considered from two points of view: the organization of their own specific form, and substances insofar as they form compounds (form and substance of expression). [TP: 43]

[the text here reads "form and content of expression" but I figure this must be a misprint?]

A stratum always has a dimension of the expressible or of expression serving as the basis for a relative invariance; for example, nucleic sequences are inseparable from a relatively invariant expression by means of which they determine the compounds, organs, and functions of the organism. [TP: 43]

Even though it is capable of invariance, expression is just as much a variable as content. [TP: 44]

To express is always to sing the glory of God. Every stratum is a judgment of God; not only do plants and animals, orchids and wasps, sing or express themselves, but so do rocks and even rivers, every stratified thing on earth. [TP: 43-4]